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Role of epistatic interactions in evolution

Project description

Exploring interactions of mutations in evolution

Mutations are the ultimate source of all genetic variation, and they may be either neutral, deleterious or beneficial in terms of their effect on the organism's ability to survive and reproduce. We understand the effects of individual mutations, but mutations may also interact with one another, a phenomenon called epistasis. Funded by the European Research Council, the EPISTAT project will study the rate and distribution of epistatic interactions among spontaneous mutations. Project findings will advance our understanding of how populations adapt to new environments and how new species arise.

Objective

The distribution of fitness effects of mutations is vital to our understanding of rates and patterns of adaptation. Population genetics and mutation accumulation experiments have given us insight into the distribution of fitness effects of single mutations. However, mutations often interact with each other. This is called epistasis. The role of epistatic interactions in adaptation has remained controversial. Yet, the distribution of epistatic effects is as fundamental as distribution of mutational effects themselves. Without knowing the distribution of epistatic effects, we cant calculate the average fitness effect of a given mutation across multiple genetic backgrounds. Moreover, if epistatic interactions tend to be positive or negative on average, this will have an important effect on evolutionary dynamics. Epistatic interactions are also known to play a role in speciation, but the proportion of mutations that exhibit incompatible interactions that can lead to speciation is unknown. I will investigate the properties of epistatic interactions among mutations with two complementary approaches. First, I will estimate the probability and distribution of effects of epistatic interactions among spontaneous mutations. I will cross mutation accumulation lines that I have developed for the fungus Neurospora crassa to produce a mapping population where spontaneous mutations are segregating, and use it to estimate the distribution of epistatic effects. Second, I will estimate the proportion of substitutions that cause reproductive incompatibilities between populations from the relationship between reproductive isolation and genetic divergence. This is achieved by an evolution experiment with fission yeast, with a design that will maximize the rate of genetic divergence with minimal change in mean phenotype. The elucidation of properties of epistatic interactions will be a major breakthrough for the field of evolutionary biology.

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HORIZON-ERC - HORIZON ERC Grants

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Call for proposal

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(opens in new window) ERC-2022-COG

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Host institution

JYVASKYLAN YLIOPISTO
Net EU contribution

Net EU financial contribution. The sum of money that the participant receives, deducted by the EU contribution to its linked third party. It considers the distribution of the EU financial contribution between direct beneficiaries of the project and other types of participants, like third-party participants.

€ 1 970 533,00
Address
SEMINAARINKATU 15
40100 Jyvaskyla
Finland

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Activity type
Higher or Secondary Education Establishments
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Total cost

The total costs incurred by this organisation to participate in the project, including direct and indirect costs. This amount is a subset of the overall project budget.

€ 1 970 533,00

Beneficiaries (1)

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