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Circuit mechanisms of cortical predictive learning

Project description

The brain’s learning mechanism for sensory predictions

Animals need to tell the difference between sensory information caused by their own movements and that from the outside world. This ability is essential for guiding behaviour, but it is a complex process. The brain learns to predict sensations generated by self-motion, using prediction errors to adjust its predictions. These errors occur when there is a mismatch between what the brain expects and what it actually senses. With this in mind, the ERC-funded Learn2Predict project explores how the brain learns from these prediction errors. By focusing on the visual cortex and the brain’s neuromodulatory system, the project aims to understand how the brain improves its sensory predictions. Using technologies like virtual reality and optogenetics, the researchers will explore how this learning process affects behaviour.

Objective

To perform sensory guided behaviors, animals need to distinguish self-generated and externally generated sensory inputs. Predictive processing theories propose that the brain does this by learning to predict sensations caused by self-motion. The key signals thought to drive this learning are prediction errors: differences between predicted and actual sensory input. My previous work shows that neurons in the primary visual cortex (V1) compute visuomotor prediction errors, and that prediction errors activate the locus coeruleus, a brain-wide neuromodulatory system. We will now investigate the circuit and neuromodulatory mechanisms underlying the learning of sensory predictions, using V1 as a model. I hypothesize that input to V1 from higher order cortical areas undergoes plasticity during self-generated sensory feedback. This plasticity should be driven by prediction errors in V1 activity, modulated by locus coeruleus output, and improve detection of externally generated visual flow during self-motion. We will test this hypothesis using innovative methods, including a multimodal virtual reality system and a novel object detection task, combined with in vivo whole cell recordings, two-photon imaging, and optogenetics. The specific aims are to (1) investigate how prediction errors are communicated between the locus coeruleus and the cortex, (2) decipher the mechanisms of predictive plasticity within the V1 circuit, and (3) assess the behavioral relevance of this plasticity. The knowledge gained will have a fundamental impact on our mechanistic understanding of predictive learning in the cortex and the role of neuromodulation in this process, which will have significance for 1) understanding conditions in which the processing of self-generated sensory feedback is thought to be disrupted (e.g. neurodevelopmental conditions and psychosis), and 2) development of AI and brain-machine interfaces that deal with self-generated sensor feedback (e.g. prostheses).

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HORIZON-ERC - HORIZON ERC Grants

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Call for proposal

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(opens in new window) ERC-2024-STG

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Host institution

THE UNIVERSITY OF EDINBURGH
Net EU contribution

Net EU financial contribution. The sum of money that the participant receives, deducted by the EU contribution to its linked third party. It considers the distribution of the EU financial contribution between direct beneficiaries of the project and other types of participants, like third-party participants.

€ 1 941 819,00
Address
OLD COLLEGE, SOUTH BRIDGE
EH8 9YL Edinburgh
United Kingdom

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Region
Scotland Eastern Scotland Edinburgh
Activity type
Higher or Secondary Education Establishments
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Total cost

The total costs incurred by this organisation to participate in the project, including direct and indirect costs. This amount is a subset of the overall project budget.

€ 1 941 819,00

Beneficiaries (1)

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