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Establishment, modulation and inheritance of sexual lineage specific DNA methylation in plants

Project description

Epigenetic inheritance in plants

Epigenetic regulation controls the expression of genes without altering the sequence of the DNA. Accumulating evidence indicates that the epigenetic status may be inherited to the next generation in both animals and plants. The EU-funded SexMeth project is interested to understand the inheritance of DNA methylation specifically in the germline of plants. Researchers will investigate the role of the environment in DNA methylation and how it may reprogramme gene expression and influence phenotype. The work will offer fundamental insight on how this epigenetic mechanism can be passed down the generations during development.

Objective

DNA methylation is an epigenetic mechanism carrying regulatory information across generations in plants and animals. Germlines called sexual lineages (SLs) in plants are essential for understanding methylation-based epigenetics because they mediate inheritance and undergo large-scale methylation reprogramming. Germline methylation reprogramming is also crucial for reproduction. However, our understanding of plant SL epigenetics is in its infancy.

I have done some of the first and most influential work in plant SL epigenetics, and developed advanced techniques for the isolation and epigenomic analysis of rare plant cell types. Recently my lab discovered that the small RNA-directed DNA methylation pathway, which generally only targets transposons, induces methylation of genes specifically in SLs, thereby regulating gene expression and meiosis. Our results also indicate that genic methylation is established in meiocytes (the origin of the male SL) by soma-derived small RNAs that are attenuated by heat stress, suggesting the hypothesis that environmentally malleable heritable information flows from soma to the germline.

We will test our hypothesis and reveal the molecular mechanisms underlying SL-specific DNA methylation in plants by pursuing the following objectives:

1. Determine how SL-specific genic methylation is established in meiocytes
2. Reveal how sRNA biogenesis and transport mediate DNA methylation in the male SL
3. Elucidate environmental modulation and transgenerational inheritance of SL methylation

Our research will reveal how new genomic loci become methylated and stay methylated through cell divisions, and how methylation is adjusted by the environment and carried to the next generation to influence phenotype. This knowledge will revolutionize our understanding of developmentally regulated methylation reprogramming, and will be invaluable for site- and/or cell type- specific engineering of DNA methylation.

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Topic(s)

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ERC-STG - Starting Grant

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Call for proposal

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(opens in new window) ERC-2018-STG

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Host institution

INSTITUTE OF SCIENCE AND TECHNOLOGY AUSTRIA
Net EU contribution

Net EU financial contribution. The sum of money that the participant receives, deducted by the EU contribution to its linked third party. It considers the distribution of the EU financial contribution between direct beneficiaries of the project and other types of participants, like third-party participants.

€ 164 728,75
Address
Am Campus 1
3400 KLOSTERNEUBURG
Austria

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Region
Ostösterreich Niederösterreich Wiener Umland/Nordteil
Activity type
Higher or Secondary Education Establishments
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Total cost

The total costs incurred by this organisation to participate in the project, including direct and indirect costs. This amount is a subset of the overall project budget.

€ 164 728,75

Beneficiaries (2)

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