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Molecular determinants of host adaptation in fungal-plant pathogens: origins and evolution of virulence effectors by genomic, phylogenetic and association mapping studies

Project description

Evolution of fungal virulence effectors

Fungal pathogens can escape plant immune defences, leading to disease emergence. Effectors are proteins secreted by fungal pathogens that play a central role in infections. Attempts to probe into the evolutionary drivers of effector diversification have been hindered by the high sequence diversity of effectors. The EU-funded EvolMAX project proposes to overcome this methodological barrier by building on the recent discovery of a structurally-conserved, but sequence-diverse family of effectors called MAX in the model fungal pathogen Magnaporthe oryzae. EvolMAX will infer the diversification history of MAX effector repertoires after host-shifts. Improved understanding of the evolutionary changes underlying host shifts will help re-engineer plants and agrosystems to durably reduce the disease burden of cereals caused by fungi.

Objective

The ability of pathogens to escape plant immune system recognition and host defenses is a significant driver of disease emergence. Small proteins secreted by fungal pathogens, named effectors, play a key role in the ability of pathogens to infect novel hosts. Fungal pathogens harbor large and highly diverse repertoires of effectors. This tremendous effector repertoire variability within and between species has hindered the characterization of effectors for their role in adaptation to novel hosts, which is limited to a tiny fraction of effectors. EvolMAX aims to lift this methodological barrier by focusing on one of the first large family of effectors discovered in fungi in order to identify the molecular processes underlying changes in virulence and adaptation to novel hosts. Building on the recent discovery of the MAX (Magnaporthe Avrs and ToxB) family of effectors in the multihost pathogen Magnaporthe oryzae, this project will jointly characterize and infer the evolutionary history of reasonably-sized sets of effectors. Using comparative genomics, population genomics, large-scale pathogenicity tests and genome-wide association mapping in host-specific M. oryzae lineages, EvolMAX will identify candidate loci involved in adaptation to novel hosts and raise understanding of the origins, diversification history of fungal effectors. The molecular processes underlying pathogen evolution and host adaptation are essential to design efficient and sustainable disease control strategies.

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MSCA-IF - Marie Skłodowska-Curie Individual Fellowships (IF)

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Call for proposal

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(opens in new window) H2020-MSCA-IF-2018

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Coordinator

INSTITUT NATIONAL DE RECHERCHE POUR L'AGRICULTURE, L'ALIMENTATION ET L'ENVIRONNEMENT
Net EU contribution

Net EU financial contribution. The sum of money that the participant receives, deducted by the EU contribution to its linked third party. It considers the distribution of the EU financial contribution between direct beneficiaries of the project and other types of participants, like third-party participants.

€ 196 707,84
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147 RUE DE L'UNIVERSITE
75007 PARIS CEDEX 07
France

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Region
Ile-de-France Ile-de-France Paris
Activity type
Research Organisations
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Total cost

The total costs incurred by this organisation to participate in the project, including direct and indirect costs. This amount is a subset of the overall project budget.

€ 196 707,84
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