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Nitric oxide-driven anaerobic oxidation of lignocellulose

Project description

Uncovering enzymes in anaerobic lignocellulose breakdown

In environments such as the riparian zones of eutrophic lakes and denitrification beds, microbes degrade lignocellulose in the absence of oxygen while also converting nitrate into nitrogen gases. This process involves complex microbial interactions and largely unknown enzymatic mechanisms. However, it remains unclear how these microbes regulate oxygen use in such anoxic environments. Additionally, enzymes that usually require oxygen for lignocellulose breakdown are present in these environments, but their exact roles remain a mystery. The ERC-funded NOD-AOL project aims to uncover how certain enzymes, including lytic polysaccharide monooxygenases (LPMOs) and nitric oxide dismutases (NODs), contribute to this process. By studying these enzymes in detail, the project will enhance our understanding of microbial carbon and nitrogen cycles.

Objective

In the riparian zone of eutrophic lakes and in field denitrification beds, microbial communities degrade lignocellulose, in anoxia, while maintaining a denitrifying lifestyle. During this process nitrate is converted to nitrogen gases via a series of reactions and intermediate products, fueled by the degradation of lignocellulosic biomass. The microbial interactions, metabolic pathways, and enzymatic mechanisms, underlining this remarkable process remain largely unknown. Intriguingly, in these anoxic habitats, enzymes involved in O2-driven oxidative lignocellulose conversion are found and expressed, yet their source of O2, their potential use of other oxidants, and their catalytic mechanism in anoxia remain unknown. Exploring this knowledge gap is a high-risk endeavor with potential to discover yet undescribed enzymes or enzyme systems capable of anaerobic oxidation of lignocellulose (AOL).

I hypothesize that oxidative cleavage of lignocellulose under denitrifying conditions follows aerobic routes including lytic polysaccharide monooxygenases (LPMOs) and lignin-active oxidases, and further, that the activity of nitric oxide dismutases (NODs) provides a source of O2 for LPMOs and lignin-active enzymes. This is conceptually high risk, linking together biological processes in a new way. Herein I will scrutinize my key hypotheses by in-depth characterization of both LPMOs, lignin-active enzymes and NODs to a depth never done before. I will study whether LPMOs can employ relevant non-conventional denitrification-linked electron acceptors and use novel approaches for characterizing the enigmatic NODs.

NOD-AOL is highly interdisciplinary and addresses questions central for understanding the global carbon and nitrogen cycles, which may ultimately help counteracting climate change.

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Topic(s)

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HORIZON-ERC - HORIZON ERC Grants

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Call for proposal

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(opens in new window) ERC-2023-COG

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Host institution

NORGES MILJO-OG BIOVITENSKAPLIGE UNIVERSITET
Net EU contribution

Net EU financial contribution. The sum of money that the participant receives, deducted by the EU contribution to its linked third party. It considers the distribution of the EU financial contribution between direct beneficiaries of the project and other types of participants, like third-party participants.

€ 1 999 858,00
Total cost

The total costs incurred by this organisation to participate in the project, including direct and indirect costs. This amount is a subset of the overall project budget.

€ 1 999 858,00

Beneficiaries (1)

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