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Elucidating the Basal Ganglia Circuits for Reward Expectation

Project description

Understanding the mechanisms of adaptive learning processes

Gaining gratification from an action or choice motivates us to make the same specific choice again. Reward expectation (RE) is basic for learning processes; the evaluation of our expectations and actual outcomes benefits our adaptive behaviour. The brain circuits of the basal ganglia nuclei are active during this process, but we still don’t know how it is generated. The EU-funded ExpectBG project aims to find in which specific circuits and cells the Reward Expectations arises and how it is computed and learned by us. Cutting-edge viral methods will be used together with electrophysiological recordings, calcium imaging techniques and specific manipulations in mice performing reinforcement learning tasks to unveil crucial mechanisms of our adaptive behaviour.

Objective

Predicting future outcomes is fundamental for adaptive behaviour. Reward-predicting stimuli evoke a state of expectation, which informs motivation, guides attention, and drives preparatory motor behaviour. Reward expectations are crucial for learning since they serve as a comparison to actual outcomes. This comparison allows animals to determine if there is a prediction error (i.e. if an outcome was better or worse than expected). Even though reward expectation signals are observed in many areas of the brain how they are computed remains unknown. The main reason for lack of progress is the absence of a clear understanding of where expectation is generated and which circuits are involved in its computation. Consequently, we are missing the prerequisite knowledge for determining where reward expectation arises, how it is computed, and how expectations are learnt. We hypothesize, based on preliminary data and prior literature, that specific circuits within the basal ganglia are crucial for computing reward expectation. We will utilize cutting edge viral methods, combined with electrophysiological recordings and calcium imaging techniques, to identify the specific circuits and cell-types within the basal ganglia nuclei that compute reward expectation. The causal role these identified circuits play in learning will be determined using cell-type specific manipulations in mice performing reinforcement learning tasks. Finally, we will pioneer approaches to manipulate elements of the basal ganglia circuit, while simultaneously recording from specific cell types in the ventral tegmental area, that are involved in computing reward prediction error. Together, this work will uncover how specific basal ganglia cell types causally contribute to the computation of reward expectation and the calculation of reward prediction error. This will provide a foundation for understating how reward expectation influences adaptive behaviour and is perturbed in psychiatric disease.

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Programme(s)

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Topic(s)

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Funding Scheme

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ERC-STG - Starting Grant

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Call for proposal

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(opens in new window) ERC-2019-STG

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Host institution

UNIVERSITY COLLEGE LONDON
Net EU contribution

Net EU financial contribution. The sum of money that the participant receives, deducted by the EU contribution to its linked third party. It considers the distribution of the EU financial contribution between direct beneficiaries of the project and other types of participants, like third-party participants.

€ 1 498 100,00
Address
GOWER STREET
WC1E 6BT LONDON
United Kingdom

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Region
London Inner London — West Camden and City of London
Activity type
Higher or Secondary Education Establishments
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Total cost

The total costs incurred by this organisation to participate in the project, including direct and indirect costs. This amount is a subset of the overall project budget.

€ 1 498 100,00

Beneficiaries (1)

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