Obiettivo The root system anchors the plant and its cells absorb water and nutrients. Since plants are sessile organisms, controlling external compound entry is essential for plant survival. In vascular plants, the endodermis is the innermost root ground tissue cell layer that controls entry to the plant vasculature by formation of a barrier for free diffusion of solutes from the soil. Moreover, many plant species also contain an exodermis layer which also acts as a barrier. The exodermis is located internal to the epidermis layer. In a differentiated state, cells of both layers contain a Casparian strip. In Arabidopsis the Casparian strip is a lignin-like structure that is deposited as a ring in the transverse section of cells and around the secondary cell wall. Recently, the developmental framework of endodermis differentiation has been described in Arabidopsis and some important molecular players identified. Here, we explore whether endodermis and exodermis differentiation are regulated similarly. Since Arabidopsis does not contain an exodermis layer, the proposed project will use the tomato root as a model system to address endodermis and exodermis differentiation at the phenotypic and molecular level. Moreover, we will address whether there are differences among species that grow in different environments similar to the environment in which their growth has been adapted. In order to address this problem, newly developed tools and technology will be used to obtain a tomato root cell-type specific transcriptome as well as data analyses required for system biology and genomic approaches. The proposed project will shed new light on endodermis and exodermis development in tomato at the phenotypic and molecular level and will lay the foundation for study in other plant species. Campo scientifico natural sciencescomputer and information sciencesdata sciencenatural sciencesbiological sciencesbiochemistrybiomoleculesproteinsagricultural sciencesagriculture, forestry, and fisheriesagriculturenatural sciencesbiological sciencesgeneticsRNAnatural sciencesearth and related environmental sciencesatmospheric sciencesclimatologyclimatic changes Programma(i) H2020-EU.1.3. - EXCELLENT SCIENCE - Marie Skłodowska-Curie Actions Main Programme H2020-EU.1.3.2. - Nurturing excellence by means of cross-border and cross-sector mobility Argomento(i) MSCA-IF-2014-GF - Marie Skłodowska-Curie Individual Fellowships (IF-GF) Invito a presentare proposte H2020-MSCA-IF-2014 Vedi altri progetti per questo bando Meccanismo di finanziamento MSCA-IF-GF - Global Fellowships Coordinatore AGENCIA ESTATAL CONSEJO SUPERIOR DE INVESTIGACIONES CIENTIFICAS Contribution nette de l'UE € 239 191,20 Indirizzo CALLE SERRANO 117 28006 Madrid Spagna Mostra sulla mappa Regione Comunidad de Madrid Comunidad de Madrid Madrid Tipo di attività Research Organisations Collegamenti Contatta l’organizzazione Opens in new window Sito web Opens in new window Partecipazione a programmi di R&I dell'UE Opens in new window Rete di collaborazione HORIZON Opens in new window Costo totale € 239 191,20 Partner (1) Classifica in ordine alfabetico Classifica per Contributo netto dell'UE Espandi tutto Riduci tutto Partner Le organizzazioni partner contribuiscono all’attuazione dell’azione, ma non sottoscrivono l’accordo di sovvenzione. THE REGENTS OF THE UNIVERSITY OF CALIFORNIA Stati Uniti Contribution nette de l'UE € 0,00 Indirizzo FRANKLIN STREET 1111 12 FLOOR 94607 OAKLAND CA Mostra sulla mappa Tipo di attività Higher or Secondary Education Establishments Collegamenti Contatta l’organizzazione Opens in new window Sito web Opens in new window Partecipazione a programmi di R&I dell'UE Opens in new window Rete di collaborazione HORIZON Opens in new window Costo totale € 160 130,40