Organisms are remarkably resilient, both in their daily life as well as during evolution. We are fascinated to know what this resilience looks like at the molecular level, both out of curiosity, but also because antibiotic or tumour drug resistance are consequences of this resilience. So how is this evolutionary resilience build in the architecture of an organisms? Because this is a very complex problem, we answer this question by focussing on the single celled organism budding yeast and on one cellular function: polarity establishment, which is essential for cell survival. Our goal is to understand, what it is in this specific network of proteins (but in any network really) that makes it so resilient during evolution. To answer this question we rebuild a simplified version of the polarity network ourselves from scratch, to observe how evolutionary resilience emerges with increasing complexity. Simultaneously we look for network properties in living cells that we can build and test in the future.